With the motion node, you can have the particles follow a path and have it oriented to it.

But when the particle goes the opposite direction it doesn’t turn around so it looks like it is going backwards. I want a rotation on the turn, like you get when you orient to path when animating on a single layer position.
My motion type is light path, I have orient to path ticked.

I\’ve fixed it, I had to orient the path of the light to \’along path\’ in its transform settings. Then my fishes started reacting to that.

Yes, since it inherits its orientation from the path when using a motion node – light path , if it isn’t oriented it has nothing to orient to. I’m guessing you had a point light, a spot should have orientation by default.

Thanks so much. This had me stumped most of the afternoon 🙂